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The genus Capoeta has a wide distribution in Southwest Asia and contains about 20 species of which 7 occur in Iran.

Its affinities are uncertain and may lie with the European Barbus/Aulopyge group or with Cyprinion and its southern and east Asian relatives (Karaman, 1971; Howes, 1982; Krupp, 1985c; Bănărescu, 1992b).

Fars, neighbourhood of Shiraz (no other locality data); CMNFI 1979-0165, 7, 30.0-96.6 mm standard length, Kerman, qanat at Ahmadabad (3032'N, 5538'E); CMNFI 1979-0166, 67, 37.1-123.1 mm standard length, Kerman, qanat at Hassanabad-e Nuq (3043'N, 5550'E); CMNFI 1979-0168, , mm standard length, Kerman, qanat at Shahabad (2907'N, 5816'E); CMNFI 1979-0169, , mm standard length, Kerman, qanat 10 km from Mahan (3008'30"N, 5717'E); CMNFI 1979-0170, , mm standard length, Kerman, qanat at Baghin (3012'N, 5648'E); CMNFI 1979-0171, , mm standard length, Kerman, qanat at Bardesir (2956'N, 5634'E); CMNFI 1979-0187, , mm standard length, Hormozgan, stream and pools at Sar Khun (2723'30"N, 5626'E); CMNFI 1979-0191, , mm standard length, Fars, stream 10 km east of Furg (ca. 5518'E); CMNFI 1979-0192, , mm standard length, Fars, qanat 2 km east of Rostaq (2826'30"N, 5504'E); CMNFI 1979-0195, , mm standard length, Fars, jube on road to Fasa (ca. 5353'30"E); CMNFI 1979-0198, , mm standard length, Fars, stream at Tadovan (2847'N, 5324'30"E); CMNFI 1979-0199, 6, 70.8-102.1 mm standard length, Fars, qanat 18 km from Jahrom (ca. 5331-40'E); CMNFI 1979-0202, , mm standard length, Fars, Mand River (2901'N, 5300'E); CMNFI 1979-0203, , mm standard length, Fars, qanat at Dudej (2933'N, 5259'E); CMNFI 1979-0204, , mm standard length, Fars, qanat on road to Kharameh (2933'N, 5259'E); CMNFI 1979-0205, 12, 45.9-200.5 mm standard length, Fars, jube at Runiz-e Pa'in (2912'N, 5342'E); CMNFI 1979-0206, , mm standard length, Fars, qanat on road to Kharameh (2912'N, 5340'E); CMNFI 1979-0207, 12, 24.2-83.7 mm standard length, Fars, jube 22 km from Neyriz (2916'N, 5428'E); CMNFI 1979-0208, 6, 39.9-130.4 mm standard length, Fars, qanat 47 km from Neyriz (ca. 5440'E); CMNFI 1979-0209, 60, 43.6-138.9 mm standard length, Kerman, qanat at Kuch Kuluh (2925'N, 5603'E); CMNFI 1979-0211, 63, 33.2-94.3 mm standard length, Kerman, river on road to Baft (2919'N, 5612'E); CMNFI 1979-0212, 73, 26.0-99.1 mm standard length, Kerman, qanat on road to Baft (2914'N, 5617'E); CMNFI 1979-0213, 5, 51.4-60.2 mm standard length, Kerman, stream in Kharan River drainage (2915'N, 5625'E); CMNFI 1979-0214, , mm standard length, Kerman, qanat pool on road to Baft (ca. 5628'E); CMNFI 1979-0215, 15, 39.7-125.9 mm standard length, Kerman, Kharan River drainage (2914'N, 5637'E); CMNFI 1979-0216, 11?

, 51.1-65.8 mm standard length, Kerman, qanat 9 km from Baft (ca. 5642'E); CMNFI 1979-0217, 15, 39.7-125.9 mm standard length, Kerman, Kharan River drainage (ca. 5651'30"E); CMNFI 1979-0221, , mm standard length, Kerman, Halil River drainage (2851'N, 5752'E); CMNFI 1979-0241, , mm standard length, Fars, Shapur River at Shapur (2947'N, 5135'E); CMNFI 1979-0243, , mm standard length, Esfahan, Zayandeh River at Falavarjan (3233'N, 5131'E); CMNFI 1979-0246, , mm standard length, Shahrestan-e Bakhtiari va Chahar Mahall, upper Karun River drainage (3157'30"N, 5059'E); CMNFI 1979-0251, , mm standard length, Esfahan, stream 1 km east of Daran (3259'N, 5026'E); CMNFI 1979-0251, , mm standard length, Esfahan, stream 1 km east of Daran (3259'N, 5026'E); CMNFI 1979-0255, , mm standard length, Markazi, Bar River drainage 2 km west of Shahabiyeh (3351'30"N, 5023'E); CMNFI 1979-0269, 1, 125.0 mm standard length, Lorestan, Dez or Karkheh River drainage (no other locality data); CMNFI 1979-0271, , mm standard length, Lorestan, Kashkan River drainage (3339'N, 4832'30"E); CMNFI 1979-0272, , mm standard length, Lorestan, river at Nokhor (3340-47'N, 4828-45'E); CMNFI 1979-0273, 7, 66.7-137.6 mm standard length, Lorestan, Kashkan River drainage (3326'N, 4819'E); CMNFI 1979-0274, 3, 28.9-141.8 mm standard length, Lorestan, Kashkan River drainage (3327'N, 4811'E); CMNFI 1979-0276, , mm standard length, Lorestan, Kashkan River drainage (ca. 4753'30"E); CMNFI 1979-0277, 2, 116.2-133.4 mm standard length, Lorestan, Kashkan River drainage (3330'N, 4759'30"E); CMNFI 1979-0278, 3, 93.5-114.7 mm standard length, Lorestan, Kashkan River drainage (3334'N, 4801'E); CMNFI 1979-0279, 1, 126.0 mm standard length, Lorestan, Khorramabad River 16 km from Nurabad (3337'N, 4818'E); CMNFI 1979-0279, 5, 115.6-155.8 mm standard length, Lorestan, Khorramabad River 16 km from Nurabad (3337'N, 4818'E); CMNFI 1979-0280, 3, 104.7-107.7 mm standard length, Lorestan, Kashkan River drainage (3343-47'N, 4812-15'E); CMNFI 1979-0282, 6, 110.3-130.3 mm standard length, Lorestan, Seymareh River drainage at Nurabad (3405'N, 4758'E); CMNFI 1979-0283, 2, 113.7-125.0 mm standard length, Kermanshahan, Qareh Su drainage (3421'N, 4707'E); CMNFI 1979-0285, 3, 125.5-148.0 mm standard length, Kermanshahan, Qareh Su drainage (3426'N, 4637'E); CMNFI 1979-0286, , mm standard length, Kermanshahan, Ravansar River at Ravansar (3443'N, 4640'E); CMNFI 1979-0287, 2, 128.2-136.1 mm standard length, Kermanshahan, Chashmeh Javari 2 km from Ravansar (ca. 4640'E); CMNFI 1979-0288, 62, 37.6-153.7 mm standard length, Ilam and Poshtkuh, Gangir River at Sarab Ewan (3350'N, 4618'E); CMNFI 1979-0289, , mm standard length, Kermanshahan, Diyala River drainage (3428'N, 4552'E); CMNFI 1979-0290, , mm standard length, Kermanshahan, Diyala River drainage at Qasr-e Shirin (3431'N, 4535'E); CMNFI 1979-0291, , mm standard length, Kermanshahan, Diyala River drainage (3424'N, 4537'E); CMNFI 1979-0306, , mm standard length, Kerman, qanat on road to Baft (2913'N, 5433'E); CMNFI 1979-0307, 5, 50.9-73.4 mm standard length, Kerman, river at Sartal 6 km from Baft (ca. 5638'E); CMNFI 1979-0308, 67, 20.5-246.9 mm standard length, Kerman, river 44 km from Baft (2902'N, 5650'E); CMNFI 1979-0309, , mm standard length, Kerman, Fahraj River at Azizabad (2857'N, 5842'E); CMNFI 1979-0315, 2, 53.5-65.5 mm standard length, Baluchestan, Bampur River 2 km from Karevandar (2751'N, 6046'E); CMNFI 1979-0315, 34?

Scaphiodon macrolepis Heckel, 1849 described from the "Confluenten des Araxes bei Persepolis" (probably the Pulvar (= Sivan) River, Fars near Persepolis) and Varicorhinus bergi Derzhavin, 1929 described in Latin from "Keredsh flumen propea Teherane, Persia septentrionalis" (Karaj River near Tehran, northern Iran) are synonyms.

Six syntypes (MNHN 2357) of Chondrostoma aculeatum in poor condition are stored in the Musum national d'Histoire naturelle, Paris (Bertin and Estve, 1948; Coad and Krupp, 1994).

Two syntypes of Scaphiodon macrolepis are in the Naturhistorisches Museum Wien under NMW 55896 and measure 100-175 mm standard length (Khsbauer, 1964).

Two other fish are marked as syntypes under NMW 51653 and are from Persepolis collected by Th. The catalogue in Vienna lists 4 fish and the 1997 card index agrees these 4 fish are the syntypes.

The extent of denticles from the base distally varies between about two-thirds and three-quarters. Scale shape is squarish with shallowly rounded to straight dorsal and ventral margins, sharp corners anteriorly, and a large to moderate central protuberance on the anterior margin.

Fars, Pulver River 8km south of Sivand (3001'30"N, 5257'E); CMNFI 1979-0061, , mm standard length, ID, more than one species?

Fars, stream tributary to Pulvar River (3004'N, 5301'E); CMNFI 1979-0063, 2, 201.0-206.7 mm standard length, Fars, qanat under Sa'adi's Tomb, Shiraz (2937'N, 5235'E); CMNFI 1979-0067, , mm standard length, Fars, qanat at Zarqan (ca. 5243'E); CMNFI 1979-0068, , mm standard length, Fars, qanat 12 km from Shiraz on Esfahan road (ca. 5234'30"E); CMNFI 1979-0073, 5, 28.9-86.6 mm standard length, Fars, Mand River beyond Chehel Chashhmeh (ca. 5201'30"E); CMNFI 1979-0074, , mm standard length, Fars, Mand River backwater (2941'N, 5206'E); CMNFI 1979-0075, , mm standard length, Fars, Mand River at Pol-e Kavar (2911'N, 5241'E); CMNFI 1979-0079, 2, 120.7-149.9 mm standard length, Fars, Mand River 5 km above Band-e Bahman (ca. 5238'E); CMNFI 1979-0079, 1, 159.7 mm standard length, Fars, Mand River 5 km above Band-e Bahman (ca. 5238'E); CMNFI 1979-0090, , mm standard length, Esfahan, Gav Khuni (ca. 5249'E); CMNFI 1979-0093, 1, 73.9 mm standard length, (); BWC 76-77 check CMNFI # CMNFI 1979-0109, 1, 91.1 mm standard length, Fars, Mand River ar Shahr-e Khafr (2856'N, 5314'E); CMNFI 1979-0109, 1, 103.4 mm standard length, Fars, Mand River ar Shahr-e Khafr (2856'N, 5314'E); CMNFI 1979-0111, 10, 8.7-54.6 mm standard length, Fars, stream 21-22 km from Shiraz (2937'30"N, 5221'E); CMNFI 1979-0113, , mm standard length, Fars, qanat under Sa'adi's Tomb (2937'N, 5235'E); CMNFI 1979-0114, , mm standard length, Fars, Mand River at road bridge (2941'N, 5206'E); CMNFI 1979-0115, 4, 154.4-172.6 mm standard length, qanat under Sa'adi's Tomb (2937'N, 5235'E); CMNFI 1979-0125, 1, 137.8 mm standard length, Bushehr, Dalaki River near Dalaki (ca. 5121'E); CMNFI 1979-0128, 16, 34.6-108.6 mm standard length, Fars, Shur River (2851'N, 5231'E); CMNFI 1979-0128, 18, 17.2-135.3 mm standard length, Fars, Shur River (2851'N, 5231'E); CMNFI 1979-0129, , mm standard length, Fars, spring 2 km north of Farrashband (2854'N, 5204'E); CMNFI 1979-0130, 5, 44.4-93.3 mm standard length, Fars, Shur River 4 km west of Firuzabad (2851'N, 5232'E); CMNFI 1979-0131, 58, 25.5-140.0 mm standard length, Fars, Mand River tributary (2838'N, 5249'E); CMNFI 1979-0132, 23, 51.1-74.4 mm standard length, Fars, Mand River tributary (2835'N, 5258'E); CMNFI 1979-0154B, 6, mm standard length, Fars, upper Shur River drainage near Darab (2845'30"N, 5224'E); CMNFI 1979-0155, 7, 36.2-80.5 mm standard length, Fars, spring at Gavanoo (2847'N, 5422'E); CMNFI 1979-0156, 3, 54.6-122.9 mm standard length, Fars, qanat at Rashidabad (2847'N, 5418'E); CMNFI 1979-0157, , mm standard length, Fars, qanat at Hadiabad (2852'N, 5413'E); CMNFI 1979-0158, , mm standard length, Fars, qanat over Qasook River (2854'N, 5353'30"E); CMNFI 1979-0159, 87, 23.1-167.3 mm standard length, Fars, qanat at Qaziabad (ca. 5343'E); CMNFI 1979-0160, 4, 66.3-138.4 mm standard length, Fars, Arteshkadeh Pomp spring (2909'N, 5337'E); CMNFI 1979-0161, 29, 33.2-88.3 mm standard length, Fars, qanat on Neyriz to Shiraz road (2910'30"N, 5341'E); CMNFI 1979-0162, 9, ?

damascina USNM 205933, 5, 97.5-142.4 mm standard length, Baluchestan, Karavandar Creek (no other locality data); ? 303854, 14, 29.1-44.6 mm standard length, Fars, Lake Arzhan (2936'N, 5159'E). damascina Capoeta aculeata(Valenciennes, 1844) Flank scale Left pharyngeal arch Common names shum (= unlucky or inauspicious, possible meaning), سياه ماهي (= siah mahi), زرده پر (= zardehpar), siah mahi aculeata.

Systematics Chondrostoma aculeatum was originally described from "eaux douces de la Perse".

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